TransactionsoftheRoyalSocietyofEdinburgh:EarthSciences,88,143-195,1998(for1997)
Articulatedthelodonts(Agnatha)ofScotland
T.MarssandA.Ritchie
ABSTRACT:Articulatedthelodonts,Loganelliascotica(Traquair),Shieliagen.n.taiti(Stetson),
LanarkiahorridaTraquair,L.spinulosaTraquairandTuriniapagei(Powrie)fromtheSilurianand
DevonianofScotlandarere-described.Anewspecies,Lanarkialanceolatasp.n.fromtheWenlock,
LowerSilurian,isestablished.Foreachspecies,diagnosisisspecified,varietiesofscalemorphology
aregiven,thebranchialareaisdescribedandbodymorphologyisdetailed.Apairedventralfinin
Shieliagen.n.taitiandLanarkialanceolatasp.n.hasbeendiscovered.Anewreconstructionof
LoganelliaandShielia,anddiagnosisforthelodontsfromScotlandaregiven.Thelocalitylistof
Scottishagnathansisreviewedandthebiostratigraphicaldistributiondiscussed.
KEYWORDS:Palaeozoicagnathans(thelodonts),morphology,squamation,biostratigraphy.
AlthoughtheSilurianthelodontshavebeenknownforoversuchareprotectedandadministeredbyScottishNatural
150years(thefirstpublicationisthatofAgassizinMurchison,Heritage.Someofthesitesarenowclosedbecauseof
1838),andarticulatedScottishspecimenswerediscoveredovercollectinganddamage.Permitstocollectnomorethan
over100yearsago(Powrie1870;Traquair1898),thebodytenspecimensfromtheothersitesmaybeobtainedfrom
st